New species of the Scirtes flavoguttatus species-group are described from SE Asia. Altogether 34 species are newly described, including Scirtes beccus sp. nov. (Malaysia), S. bocakorum sp. nov. (Indonesia), S. crockerensis sp. nov. (Malaysia), S. decorus sp. nov. (Malaysia), S. dumogensis sp. nov. (Indonesia), S. gunongmulensis sp. nov. (Malaysia), S. ishikawai sp. nov. (Vietnam), S. kinabalensis sp. nov. (Malaysia), S. kundasangensis sp. nov. (Malaysia), S. lambriensis sp. nov. (Indonesia), S. leuserensis sp. nov. (Indonesia), S. luteus sp. nov. (Malaysia), S. malaisei sp. nov. (Myanmar), S. melinauensis sp. nov. (Malaysia), S. noonadan sp. nov. (Philippines), S. pallicolor sp. nov. (Malaysia), S. penampangensis sp. nov. (Malaysia), S. phoupanensis sp. nov. (Laos), S. prodigiosus sp. nov. (Malaysia), S. punctatus sp. nov. (Philippines), S. quasibalehensis sp. nov. (Malaysia), S. ranauensis sp. nov. (Malaysia), S. sarawakensis sp. nov. (Malaysia), S. seblatensis sp. nov. (Indonesia), S. sibayensis sp. nov. (Indonesia), S. sibolangitensis sp. nov. (Indonesia), S. sulawesicus sp. nov. (Indonesia), S. sulcigeroides sp. nov. (Malaysia), S. talinisensis sp. nov. (Philippines), S. ulukimanisensis sp. nov. (Malaysia), S. velutinus sp. nov. (Malaysia), S. vietnamicus sp. nov. (Vietnam), S. wallacei sp. nov. (Indonesia), S. yangsinensis sp. nov. (Vietnam). New localities of six species are provided. An updated identification key, checklist and a summary of distributional data are included. Probability of the occurrence of the Scirtes flavoguttatus species-group was evaluated with an analysis in MaxEnt software. It is highly plausible that members of the group occur in most mountainous rainforests of SE Asia.
Matched MeSH terms: Beetles/anatomy & histology; Beetles/classification*; Beetles/growth & development
The Oriental pselaphine genus Horniella Raffray, 1905 (tribe Tyrini: subtribe Somatipionina) is redefined and revised. Twenty-five new species are described: H. centralis Yin & Li, sp. n., H. confragosa Yin & Li, sp. n., H. dao Yin & Li, sp. n., H. hongkongensis Yin & Li, sp. n., H. nakhi Yin & Li, sp. n., H. schuelkei Yin & Li, sp. n., H. sichuanica Yin & Li, sp. n., H. simplaria Yin & Li, sp. n., and H. tianmuensis Yin & Li, sp. n. from China, H. himalayica Yin & Li, sp. n. from Nepal and North India, H. asymmetrica Yin & Li, sp. n., H. burckhardti Yin & Li, sp. n., H. intricata Yin & Li, sp. n., H. kaengkrachan Yin & Li, sp. n., H. khaosabap Yin & Li, sp. n., H. loebli Yin & Li, sp. n., H. phuphaman Yin & Li, sp. n., H. prolixo Yin & Li, sp. n., and H. schwendingeri Yin & Li, sp. n. from Thailand, H. philippina Yin & Li, sp. n. from the Philippines, H. awana Yin & Li, sp. n., H. gigas Yin & Li, sp. n., H. pilosa Yin & Li, sp. n., and H. smetanai Yin & Li, sp. n. from Malaysia, and H. cibodas Yin & Li, sp. n. from Indonesia. The two previously described species, H. hirtella Raffray, 1901 (type species) from Sri Lanka and H. falcis Yin & Li, 2010 from China are redescribed, and a lectotype is designated for H. hirtella. Illustrations of habitus and important diagnostic features, an identification key, and distributional maps for all species are provided. Eleven unidentified species represented only by females are left unnamed. Illustrations of the habitus and the genital complex, and label data of these species are given to facilitate future study. All available data indicates that species of Horniella typically inhabit leaf litter of various kinds of forests, and can be most efficiently collected by sifting and use of Winkler-Moczarski extractors.
Matched MeSH terms: Beetles/anatomy & histology; Beetles/classification*; Beetles/growth & development
A new species, Megatyrus femoralis sp. n., is described from the Koshi Zone, East Nepal, with major diagnostic features illustrated. Megatyrus masumotoi Nomura, Sakchoowong & Chanpaisaeng, originally described from southwestern Thailand, is recorded from the Noring Timur Mountain, West Malaysia. The above data extends the known range of Megatyrus about 1,200 km to the west, and 870 km to the south.
The Pediacus Shuckard fauna of Asia and Australasia is revised. Eighteen species are recorded, described and illustrated from the regions and a key to species is provided. Nine new species are described: Pediacus australis sp. nov. (Australia, Papua New Guinea, Philippines, Thailand), P. carinatus sp. nov. (Indonesia, Malaysia, Thailand), P. fujianensis sp. nov. (China), P. japonicoides sp. nov. (Taiwan); P. leei sp. nov. (Taiwan), P. pendleburyi sp. nov. (Malaysia), P. sinensis sp. nov. (China), P. taiwanensis sp. nov. (Taiwan) and P. thomasi sp. nov. (Taiwan). A checklist of the Pediacus fauna of the world is given, listing a total of 31 species.
Two new species of Ancyronyx Erichson, 1847 (Coleoptera: Elmidae) are described from Borneo: A. pulcherrimus (Brunei) and A. reticulatus (Sabah). Habitus views, illustrations of important characters as well as plastron structures of Ancyronyx reticulatus are presented and discussed.
Pycnotarsobrentus inuiae Maruyama & Bartolozzi, gen. nov. and sp. nov. (Brentinae: Eremoxenini) is described from the Lambir Hills National Park, Borneo (Sarawak, Malaysia) based on specimens collected from Crematogaster difformis F. Smith, 1857 ant nests in the myrmecophytic epiphytic ferns Platycerium crustacea Copel. and Lecanopteris ridleyi H. Christ. A second species of Pycnotarsobrentus is known from Malaysia but is represented by only one female and consequently not yet described pending discovery of a male. Pycnotarsobrentus belongs to the tribe Eremoxenini and shares some character states with the African genus Pericordus Kolbe, 1883. No species of Eremoxenini with similar morphological modifications are known from the Oriental region.
Matched MeSH terms: Beetles/anatomy & histology; Beetles/classification*; Beetles/growth & development
Anew hydrophilid genus Chimaerocyon gen. nov. containing two species, C. shimadai sp. nov. (Malaysia: Pahang) and C. sumatranus sp. nov. (Indonesia: Sumatra), is described. Specimens of C. shimadai were collected from brood cells in anest of Pheidole singaporensis Özdikmen, 2010. The biology of C. sumatranus remains unknown. A molecular phylogeny based on four genes (cox1, cox2, 18S and 28S) supports the placement of the genus as deeply nested within the Cercyon-group of the tribe Megasternini. This position is supported by the subdistal position of the median spur in the hind wing (unique to Megasternini) and the presence of sucking disc on male maxilla (unique for Megastemini+Sphaeridiini). The remaining external morphology differs substantially from other representatives of Megasternini. The hypothesis that the aberrant morphology of Chimaerocyon gen. nov. is a consequence of myrmecophily is discussed.
Three species have been recognized in the genus Geoparnus Besuchet, 1978, all collected by sifting ground debris from tropical rainforests of Malaysia. The new species Geoparnus loebli sp. nov. is described; morphological similarities to known species G setifer Besuchet, 1978 and G rhinoceros Kodada, Jach, Ciampor Jr & (Camporova-Zat'ovicova, 2007 are discussed. Habitus of adult, male and female genitalia as well as other diagnostic characters are illustrated.
Five new species of Notosacantha are described from Borneo (Sabah): N. flaviventris Borowiec and Takizawa sp. nov. (also described from Sumatra), N. flavosuturata Świętojańska and Takizawa sp. nov., N. minuta Świętojańska and Takizawa sp. nov., N. obscuricornis Borowiec and Takizawa sp. nov. and N. ovoidea Borowiec and Świętojańska sp. nov. New records for nine other species, a check list and key to Bornean Notosacantha are given. Myrsinaceae and Myristicaceae are new host plant families for tortoise beetles and Ardisia elliptica Thunb., Ardisia sp., Embelia sp., Gironiema sp. (all Myrsinaceae) and Knema sp. (Myristicaceae) are new host plants for Notosacantha.
Tmesiphodimerus Coulon and Yin, new genus (Pselaphitae: Tmesiphorini) is proposed for T. sinensis Yin and Coulon, new species from Hainan, South China (type species), and T. malaysianus Coulon and Yin, new species from Perak, West Malaysia. The new taxa are described, with their major diagnostic features illustrated. The taxonomic placement of Tmesiphodimerus is discussed.
Matched MeSH terms: Beetles/anatomy & histology; Beetles/classification*; Beetles/growth & development
Motschulsky (1859), a Russian Imperial Army Colonel and entomologist, established the genus Exopholis, described E. birmannica and transferred Melolontha hypoleuca Wiedemann, 1819 to it. Brenske (1896) described Exopholis philippinica from the "Philippinen" from a single female specimen. Dalla Torre (1912), listed eight species under the genus Exopholis Motschulsky, 1859 from South East Asia: E. hypoleuca (Weidemann, 1819) (Myanmar, Malaysia, Borneo, Java, Sumatra, Nias, Ambon), E. costata (Burmeister, 1855) (Java, Nias), E. birmanica Motschulsky, 1859 (Myanmar), E. lacordairei Waterhouse, 1867 (Borneo), E. pinguis Lansberge, 1879 (Sumatra), E. brenskei Nonfried, 1891, E. borneensis Brenske, 1894 (both from Borneo) and E. philippinica Brenske, 1896 (Philippines). This paper aims to review the taxonomy of Exopholis from the Philippines. Specifically, to focus on the diagnosis of Exopholis philippinica and its geographic distribution.
Nipponophloeostiba thayeri Shavrin, sp.n. from Borneo Island, Sabah (Malaysia) is described, illustrated and compared with Japanese N. verrucifera Watanabe, 1962.
A new species of Clidicini ant-like stone beetles, Clidicus mawarensis sp. n., is described and illustrated. The holotype male was collected in East Malaysia (Borneo: Sabah); the new species belongs to a group of large-bodied Clidicus, and shows similarities to C. ganglbaueri Reitter; the male has unusually complex structures of the aedeagal apical region.
The genus Airapus Stebnicka Howden, 1996 currently comprises 26 extant species distributed in the Australian and Oriental zoogeographical regions (Stebnicka Howden 1996; Stebnicka 1998, 2009; Rakovič et al. 2019; Král et al. 2019; Minkina 2020) and one fossil species from the Eocene Baltic amber (Tamutis et al. 2017). Of the continental Southeast Asia, only three species have been known so far: Airapus cechovskyi Král, Mencl Rakovič, 2019 (mainland Malaysia: Kelantan), A. tyri Král, Mencl Rakovič, 2019 (Central Thailand: Phetchaburi Province) and A. sicardi ( Paulian, 1945) (Laos: "Cochinchine: Long Xuyen" and South Vietnam: "Annam: Tanh Hoa") (Paulian 1945; Balthasar 1964; Král et al. 2019). Examination of the material housed in the collections of the Institute of Zoology, Chinese Academy of Sciences, Beijing, China, revealed Airapus material belonging to an undescribed species. Its formal description is presented in this paper. This new species is another, fourth species occurring in mainland Southeast Asia. It is also the first country record from China. The geographical distribution of the genus is now known to the north as far as Fujian Province.
Two new species of the genus Odochilus Harold, 1877 (Coleoptera: Scarabaeidae: Aphodiinae: Odochilini) from the Oriental Region are described as follows: O. shavrini Rakovič Anichtchenko, new species from the Philippines (Mindanao) and O. borneensis Rakovič Anichtchenko, new species from Malaysia (Sarawak). Photographs of their habitus are presented. Differential diagnoses are mentioned.
Based on material collected by canopy fogging in forests on Borneo, Obhylius arboricola gen. nov., sp. nov., and Seticotasteromimus brunomanseri sp. nov., the second species of this genus, are described. Further records of the type species of Seticotasteromimus Germann, 2013, S. jarawa Germann, 2013, are presented from Borneo, some 2700 km distant from the type locality on the Andaman Islands. Along with Seticotasteromimus, the new genus is best placed in the subtribe Cotasteromimina.
The monotypic genus Paracyriothasastes Breuning, 1978 was established for Cereopsius marmoreus Pascoe, 1857 from Malaysia. Uraechoides Breuning, 1981 was established for Uraechoides vivesi Breuning, 1981 also from Malaysia, and is currently composed of the type species and U. taomeiae Hayashi, Nara Yu, 1995, the latter from China (Taiwan) (Tavakilian Chevillotte 2020).
Six new species of the cephenniine genus Hlavaciellus Jałoszyński are described: H. cincinnalis sp. n. (Malaysia: Sabah); H. carinatus sp. n. (Malaysia: Pahang); H. microtuberculatus sp. n. (Malaysia: Pahang); H. diversipilosus sp. n. (Malaysia: Pahang); H. concavus sp. n. (Indonesia: Sumatra); and H. sumatranus sp. n. (Indonesia: Sumatra). An updated key to identification of males of all nominal species of Hlavaciellus is given.
A new genus and five new species of neotenic lyropaeine net-winged beetles, Plycus prosvirovi gen. nov., sp. nov., Platerodrilus (s. str.) brastagicola sp. nov., P. (s. str.) gorbunovi sp. nov., P. (s. str.) kambanganus sp. nov. and P. (Duliticola) berastagiensis sp. nov., are described from Malaysia and Indonesia. Horakiella Bocakova, 2006, syn. nov. is proposed as a junior synonym of Atamania Kazantsev, 2005. Descriptions of little known members of the subfamily, Lybnopaeus avernus (Kazantsev, 1997), Lyropaeus (Lyroneces) orcinus Kazantsev, 1998, Miniduliticola nelsoni Kazantsev, 2002, Platerodrilus (Zhelongia) grootaerti Kazantsev, 2009, P. (Zh.) holynskae Kazantsev, 2009, P. (Duliticola) svetae Kazantsev, 2009, P. (D.) strbai Kazantsev, 2009, Atamania tembeling Kazantsev, 2002 and Sinodulia gressitti Kazantsev, 2002, are complemented with macrophotographs. Lists of Platerodrilus and Atamania species are provided.
The genus Morphosphaera Baly, 1861 is revised. Eleven species are considered as valid, including, M. takizawai sp. nov. (Mt. Basor, 90 km N of Gua Musang, Malaysia, W. Kelantan), described from Malaysia and Indonesia. Color photos of habitus and drawings of diagnostic characters from eleven species are presented. The following synonymies are proposed: M. sodalis Chen, 1935 and M. brunnea Maulik, 1936 are junior synonyms of M. albipennis Allard, 1889; M. margaritacea Laboissière, 1930, M. viridipennis Laboissière, 1930, and M. prava Maulik, 1936 are junior synonyms of M. coomani Laboissière, 1930; M. gracilicornis Chen, 1963 is a junior synonym of M. maculicollis Baly, 1861; M. cavaleriei Laboissière, 1930, M. cincticollis Laboissière, 1930, M. marginata Laboissière, 1930, M. purpurea Laboissière, 1930, M. gingkoae Gressitt & Kimoto, 1963, and M. metallescens Gressitt & Kimoto, 1963 are junior synonyms of M. sumatrana Jacoby, 1886. The type material of M. impunctata Allard, 1890 from the Philippines was not found and its taxonomic status remains uncertain. Morphosphaera peregrina Weise, 1913 is transferred to the genus Borneola Mohamedsaid, 1998 nov. comb. A neotype is designated for Chrysomela japonica Hornstedt, 1788. Lectotypes are designated for the following species: Adorium chrysomeloides Bates, 1866, A. japonicum Baly, 1874, Morphosphaera albipennis Allard, 1889, M. bimaculata Chûjô, 1938, M. caerulea Jacoby, 1896, M. cavaleriei Laboissière, 1930, M. collaris Laboissière, 1930, M. formosa Laboissière, 1930, M. marginata Laboissière, 1930, M. montivaga Maulik, 1936, M. prava Maulik, 1936, M. purpurea Laboissière, 1930, M. sumatrana Jacoby, 1886, M. viridipennis Laboissière, 1930, and Galerucida simplex Weise, 1922.