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  1. Ramiah SK, Awad EA, Mookiah S, Idrus Z
    Poult Sci, 2019 Sep 01;98(9):3828-3838.
    PMID: 30839088 DOI: 10.3382/ps/pez093
    The use of nanominerals, such as nano-zinc, represents a promising and emerging technology in the animal farming industry. Due to the small particle size and bioavailability of nano-zinc, it can be easily assimilated in the digestive system, thereby reducing excretion and environmental pollution. The present study was conducted to assess the effects of zinc oxide nanoparticles (ZnONPs) on the growth performance, zinc (Zn) concentration in edible tissues, thiobarbituric acid reactive substance, and corticosterone concentrations in broilers reared under normal or heat stress environmental conditions. The experiment was performed with a completely randomized design based on a 4 × 2 factorial arrangement consisting of 4 diets (basal diet + 60 mg/kg conventional zinc oxide as control diet; basal diet + 40 mg/kg of ZnONPs; basal diet + 60 mg/kg of ZnONPs; and basal diet + 100 mg/kg of ZnONPs) and 2 environmental conditions (normal and heat stress). On day 22, birds from each dietary group were divided equally to normal temperature (23 ± 1°C throughout) or heat stress conditions (34 ± 1°C daily for 6 h from 10:00 am until 4:00 pm). From 1 to 42 D of age, the broiler chickens fed 100 mg/kg ZnONPs exhibited lower feed intake and feed conversion ratio than the control. The accumulation of Zn in the liver of broilers was significantly higher among all treatment groups compared to breast and thigh muscle tissues regardless of the temperature conditions. At 40 and 60 mg/kg ZnONPs, the malondialdehyde content increased in thigh muscle of broilers at 7 D postmortem, indicating that ZnONPs potentially inhibited the antioxidant system in muscle tissues. The control and ZnONPs at 40 mg/kg and 60 mg/kg led to low serum corticosterone levels that may be attributed to the antioxidant and antistress properties of Zn. Taken together, although supplementation with ZnONPs at 40 mg/kg and 60 mg/kg alleviated the negative results of heat stress, further research is needed to determine the optimal level of dietary ZnONPs supplementation.
  2. Najafi P, Zulkifli I, Jajuli NA, Farjam AS, Ramiah SK, Amir AA, et al.
    Int J Biometeorol, 2015 Nov;59(11):1577-83.
    PMID: 25649005 DOI: 10.1007/s00484-015-0964-3
    An experiment was conducted to determine the effect of different stocking densities on serum corticosterone (CORT), ovotransferrin (OVT), α1-acid glycoprotein (AGP) and ceruloplasmin (CP) concentrations, brain heat shock protein (HSP) 70 expression and performance in broiler chickens exposed to unheated and heated conditions. Day-old chicks were stocked at 0.100 m(2)/bird (low density (LD)) or 0.063 m(2)/bird (high density (HD)), in battery cages and housed in environmentally controlled rooms. From 21 to 35 days of age, birds from each stocking density group were exposed to either 24 or 32 °C. Growth performance was recorded during the heat treatment period, and blood and brain samples were collected to determine CORT, OVT, AGP, CP and HSP 70 levels on day 35. Heat treatment but not stocking density was detrimental to growth performance. There were significant temperature × density interactions for CORT, CP and OVT on day 35. Although HD elevated CORT, CP and OVT when compared to LD, the effects of the former were more obvious under heated condition. Both temperature and density had significant effect on AGP and HSP 70. In conclusion, irrespective of temperature, high stocking density was physiologically stressful to broiler chickens, as indicated by CORT, AGP, CP, OVT and HSP 70, but not detrimental to growth performance and survivability. As it was shown in the present study, AGP, CP and OVT could be useful biomarkers to determine the effect of overcrowding and high temperature on the welfare of broiler chickens.
  3. Ramiah SK, Meng GY, Sheau Wei T, Swee Keong Y, Ebrahimi M
    PPAR Res, 2014;2014:137652.
    PMID: 25309587 DOI: 10.1155/2014/137652
    Conjugated linoleic acids (CLA) act as an important ligand for nuclear receptors in adipogenesis and fat deposition in mammals and avian species. This study aimed to determine whether similar effects are plausible on avian abdominal fat adipocyte size, as well as abdominal adipogenic transcriptional level. CLA was supplemented at different levels, namely, (i) basal diet without CLA (5% palm oil) (CON), (ii) basal diet with 2.5% CLA and 2.5% palm oil (LCLA), and (iii) basal diet with 5% CLA (HCLA).The content of cis-9, trans-11 CLA was between 1.69- and 2.3-fold greater (P < 0.05) than that of trans-10, cis-12 CLA in the abdominal fat of the LCLA and HCLA group. The adipogenic capacity of the abdominal fat depot in LCLA and HCLA fed chicken is associated with a decreased proportion of adipose cells and monounsaturated fatty acids (MUFA). The transcriptional level of adipocyte protein (aP2) and peroxisome proliferator-activated receptor gamma (PPARγ) was downregulated by 1.08- to 2.5-fold in CLA supplemented diets, respectively. It was speculated that feeding CLA to broiler chickens reduced adipocyte size and downregulated PPARγ and aP2 that control adipocyte cellularity. Elevation of CLA isomers into their adipose tissue provides a potential CLA-rich source for human consumption.
  4. Saminathan M, Mohamed WNW, Noh 'M, Ibrahim NA, Fuat MA, Ramiah SK
    Trop Anim Health Prod, 2022 Jan 17;54(1):64.
    PMID: 35038035 DOI: 10.1007/s11250-022-03046-5
    Palm oil is a natural energy source ingredient in poultry diets that offers a broad range of beneficial effects on the performance of broiler chickens. This review was conducted to highlight the impact of palm oil as a feed ingredient on growth performance and carcass quality, as well as the biochemical, antioxidant activity and tissue fatty acids (FA) composition of broiler chickens. Palm oil inclusion in broiler chickens' rations contributes significantly to the high metabolisable energy (ME) of feed formulation, increases feed palatability and decreases digesta passage rate in the intestine. The reviewed literature indicated that dietary palm oil has a beneficial effect on broiler chickens' overall growth performance traits. The addition of palm oil can also improve the heat tolerance of chickens reared in high ambient temperature conditions. Regardless of breed and breeding conditions, palm oil exhibits good oxidative stability in broiler chickens due to the presence of prevalent phytonutrient elements in this oil. The inclusion of palm oil increased palmitic (C16:0) and oleic (C18:1) acids in tissue deposits, which improves meat stability and quality. Moreover, molecular studies have revealed that higher mRNA expression of several lipid-related hepatic genes in broiler chickens fed palm oil. Nonetheless, dietary palm oil can influence FA deposition in tissues, modulate lipoprotein and triglycerides (TG) levels, and cytokine contents in the blood serum of broiler chickens.
  5. Ramiah SK, Balakrishnan KN, Subramaniam Y, Iyasere OS, Idrus Z
    Animals (Basel), 2022 Nov 29;12(23).
    PMID: 36496875 DOI: 10.3390/ani12233354
    The phenomenon of increasing heat stress (HS) among animals is of particular significance when it is seen in economically significant industries, such as poultry. Due to the identification of the physiological, molecular, and genetic roots of HS responses in chickens, a substantial number of studies have focused on reducing the effects of HS in poultry through environmental management, dietary manipulation, and genetic alterations. Temperature manipulation (TM) during embryogenesis has been claimed to increase the thermal tolerance and well-being of chickens without affecting their capacity for future growth. There has been little investigation into the vulnerability of the epigenome involving TM during embryogenesis, although the cellular pathways activated by HS have been explored in chickens. Epigenetic changes caused by prenatal TM enhance postnatal temperature adaption and produce physiological memory. This work offers a thorough analysis that explains the cumulative impact of HS response genes, such as genes related to heat shock proteins, antioxidants, and immunological genes, which may aid in the enhanced adaptability of chickens that have undergone thermal manipulation during their embryonic stages.
  6. Hanafi SA, Zulkifli I, Ramiah SK, Chung ELT, Kamil R, Awad EA
    Poult Sci, 2023 Feb;102(2):102390.
    PMID: 36608455 DOI: 10.1016/j.psj.2022.102390
    Prenatal stress may evoke considerable physiological consequences on the developing poultry embryos and neonates. The present study aimed to determine prenatal auditory stimulation effects on serum levels of ceruloplasmin (CPN), alpha-1-acid glycoprotein (AGP), corticosterone (CORT), and heat shock protein 70 (Hsp70) regulations in developing chicken embryos and newly hatched chicks. Hatching eggs were subjected to the following auditory treatments; 1) control (no additional sound treatment other than the background sound of the incubator's compressors at 40 dB), 2) noise exposure (eggs were exposed to pre-recorded traffic noise at 90 dB) (NOISE), and 3) music exposure (eggs were exposed to Mozart's Sonata for Two Pianos in D Major, K 488 at 90 dB) (MUSIC). The NOISE and MUSIC treatments were for 20 min/h for 24 h (a total of 8 h/d), starting from embryonic days (ED) 12 to hatching. The MUSIC (1.37 ± 0.1 ng/mL) and NOISE (1.49 ± 0.2 ng/mL) treatments significantly elevated CPN at ED 15 compared to the Control (0.82 ± 0.04 ng/mL) group and post-hatch day 1 (Control, 1.86 ± 0.2 ng/mL; MUSIC, 2.84 ± 0.4 ng/mL; NOISE, 3.04 ± 0.3 ng/mL), AGP at ED 15 (Control, 39.1 ± 7.1 mg/mL; MUSIC, 85.5 ± 12.9 mg/mL; NOISE, 85.4 ± 15.1 mg/mL) and post-hatch day 1 (Control, 20.4 ± 2.2 mg/mL; MUSIC, 30.5 ± 4.7 mg/mL; NOISE, 30.3 ± 1.4 mg/mL). CORT significantly increased at ED 15 in both MUSIC (9.024 ± 1.4 ng/mL) and NOISE (12.15 ± 1.6 ng/mL) compared to the Control (4.39 ± 0.7 ng/mL) group. On the other hand, MUSIC exposed embryos had significantly higher Hsp70 expression than their Control and NOISE counterparts at ED 18 (Control, 12.9 ± 1.2 ng/mL; MUSIC, 129.6 ± 26.4 ng/mL; NOISE, 13.3 ± 2.3 ng/mL) and post-hatch day 1 (Control, 15.2 ± 1.7 ng/mL; MUSIC, 195.5 ± 68.5 ng/mL; NOISE, 13.2 ± 2.7 ng/mL). In conclusion, developing chicken embryos respond to auditory stimulation by altering CPN, AGP, CORT, and Hsp70. The alterations of these analytes could be important in developing embryos and newly hatched chicks to cope with stress attributed to auditory stimulation.
  7. Ramiah SK, Zulkifli I, Rahim NA, Ebrahimi M, Meng GY
    Asian-Australas J Anim Sci, 2014 Mar;27(3):375-82.
    PMID: 25049964 DOI: 10.5713/ajas.2013.13030
    The competency of garlic and pennywort to improve broiler chicken growth and influence intestinal microbial communities and fatty acid composition of breast meat were studied. Two hundred forty, "day-old" chicks were randomly allocated to 4 treatment groups consisting of 6 replications of 10 chicks in each pen. The groups were assigned to receive treatment diets as follows: i) basal diet (control), ii) basal diet plus 0.5% garlic powder (GP), iii) basal diet plus 0.5% pennywort powder (PW) and iv) 0.002% virginiamycin (VM). Birds were killed at day 42 and intestinal samples were collected to assess for Lactobacillus and Escherichia coli. The pectoralis profundus from chicken breast samples was obtained from 10 birds from each treatment group on day 42 and frozen at -20°C for further analyses. Fatty acid profile of breast muscles was determined using gas liquid chromatography. Feed intake and weight gain of broilers fed with GP, PW, and VM were significantly higher (p<0.05) compared to control. Feeding chicks GP, PW, and VM significantly reduced Escherichia coli count (p<0.05) while Lactobacillus spp count were significantly higher (p<0.05) in the gut when compared to control group on day 42. Supplemented diet containing pennywort increased the C18:3n-3 fatty acid composition of chickens' breast muscle. Garlic and pennywort may be useful in modulating broiler guts as they control the enteropathogens that help to utilize feed efficiently. This subsequently enhances the growth performances of broiler chickens.
  8. Ahmad-Hanafi S, Zulkifli I, Ramiah SK, Chung ELT, Kamil R, Sazili AQ, et al.
    Poult Sci, 2024 Oct;103(10):103948.
    PMID: 39127008 DOI: 10.1016/j.psj.2024.103948
    Feed restriction could induce physiological stress in broiler chickens, leading to welfare issues. Prenatal stimulation could improve stress-coping mechanisms in poultry. The present study aimed to elucidate the effects of subjecting developing embryos to auditory stimulation on physiological stress response to feed restriction in broiler chickens at market age. A total of 423 hatching eggs of Cobb 500 (Gallus domesticus) were subjected to the following auditory treatments: 1) no additional sound treatment other than the background sound of the incubator's compressors at 40 dB (CONTROL), 2) exposure to pre-recorded traffic noise at 90 dB (NOISE), and 3) exposure to Mozart's Sonata for Two Pianos in D Major, K 488 at 90 dB) (MUSIC). The NOISE and MUSIC treatments were for 20 min/h for 24 h (a total of 8 h/d), starting from embryonic days (ED) 12 to hatching. On d 42, an equal number of birds from each prenatal auditory stimulation (PAS) group were subjected to either ad libitum feeding (AL) or 30-h of feed restriction (FR) in a completely randomised design. The FR chickens exhibited significantly higher serum levels of corticosterone (CORT), and heat shock protein (HSP) 70 compared to those of AL. Prenatal auditory stimulation, particularly NOISE, led to lower serum levels of CORT and alpha-1-acid glycoprotein (AGP) levels compared to the CONTROL group. Additionally, NOISE significantly increased brain mRNA glucocorticoid receptor and HSP70 gene expression. The cecal population of E. coli and Lactobacillus spp. was not significantly affected by prenatal auditory stimulation. In conclusion, our findings suggest that prenatal auditory stimulation, particularly NOISE, positively impacts broiler chickens' ability to cope with feed restriction.
  9. Ramiah SK, Atta Awad E, Hemly NIM, Ebrahimi M, Joshua O, Jamshed M, et al.
    J Anim Sci, 2020 Oct 01;98(10).
    PMID: 32936879 DOI: 10.1093/jas/skaa300
    This study was conducted to explore the effect of the zinc oxide nanoparticles (ZnONPs) supplement on the regulatory appetite and heat stress (HS) genes in broiler chickens raised under high or normal ambient temperatures. In this study, 240 one-day-old male broiler chicks (Cobb 500) were randomly assigned to 48 battery cages. From day 1, these 48 cages were randomly subjected to four different treatment strategies: Control (wherein, their basal diet included 60 mg/kg of ZnO), ZNONPs 40 (wherein basal diet included 40 mg/kg of ZnONPs), ZnONPs 60 (basal diet included 60 mg/kg of ZnONPs), and ZnONPs 100 (basal diet included 100 mg/kg of ZnONPs). Thereafter, from day 22 to 42, the chickens from each dietary treatment group were subjected to different temperature stresses either normal (23 ± 1 °C constant) or HS (34 ± 1 °C for 6 h/d), which divided them into eight different treatment groups. Our findings revealed that dietary ZnONPs altered the gene expression of cholecystokinin (ileum), heat stress proteins (HSP) 70 (jejunum and ileum), and HSP 90 (duodenum, jejunum, and ileum). The gene expression of ghrelin was affected by the interaction between the ZnONPs concentration and temperature in the duodenum and stomach. More studies are required to elucidate its complex physiological and biochemical functions of the regulation of gene expression within the intestine in heat-stressed broiler chickens.
  10. Saminathan M, Gan HM, Abdullah N, Wong CMVL, Ramiah SK, Tan HY, et al.
    J Appl Microbiol, 2017 Apr 23.
    PMID: 28434189 DOI: 10.1111/jam.13477
    AIMS: To evaluate the effects of condensed tannins (CTs) fractions of differing molecular weights (MWs) from a Leucaena leucocephala hybrid-Rendang on the rumen protozoal community in vitro.

    METHODS AND RESULTS: The effects of unfractionated CTs (F0) and CT fractions of different MWs (F1 > F2 > F3 > F4 > F5) on protozoal population and community were evaluated in vitro using rumen microbes and ground guinea grass as the substrate. Higher-MW CT fractions F1 and F2 significantly (P 

  11. Mookiah S, Wan Mohamed WN, Md Noh ', Ibrahim NA, Fuat MA, Ramiah SK, et al.
    PMID: 32898954 DOI: 10.5713/ajas.20.0360
    Ahead of Print article withdrawn by publisher.
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