The ground beetle genus Hexachaetus Chaudoir, 1871 is re-defined and reviewed. Bearing six setae on ligula is no more considered as a crucial characteristic for Hexachaetus. Members of Hexachaetus share the following combination of morphological features: body polish, smooth, and impunctate, ligula more or less dilated at apex, bearing 4, 6, or even 12 setae apically, prosternal process unbordered at apex, elytra distinctly and obliquely truncated at apex, with the apical inner angles very sharp in most species (except for H. mulan n. sp.), and interval 3 with anterior and posterior setiferous pores (median one lacking). The members of Hexachaetus are about 20 species which could be divided into six species groups. All except angulatus species group are dealt with in this paper, with descriptions of four new species: H. kirschenhoferi n. sp. (Indonesia: Kalimantan), H. brunki n. sp. (Malaysia: N. Borneo), H. vietnamensis n. sp. (Vietnam: Annam) and H. mulan n. sp. (Malaysia: Perak and Pahang). H. maindroni Tian & Deuve, 2006 is proposed as a subspecies of H. lateralis Guérin, 1843, n. stat. A key to species groups and species of the genus is also provided.
Matched MeSH terms: Beetles/anatomy & histology; Beetles/classification*; Beetles/growth & development
A new species of riffle beetle, Vietelmis jablonskii, from Laos and Thailand is described and illustrated. Differences to other Vietelmis species are discussed and an identification key for the Vietelmis is presented. New faunistic records are provided for V. kovaci Kodada & Čiampor, 2000; the genus Vietelmis is newly recorded for Thailand.
The genus Doryscus Jacoby, 1887 is revised. Twelve new species are described: D. indochinensis sp. nov. from China (Yunnan), India, Laos, Thailand, Vietnam; D. krausi sp. nov. from Laos and Thailand; D. kubani sp. nov., from China (Yunnan); D. geiseri sp. nov. from Singapore and Thailand; D. nepalensis sp. nov. from Bhutan, north India, and Nepal; D. luzonensis sp. nov., and D. mindanaoensis sp. nov. from Philippines; D. barclayi sp. nov., D. boreri sp. nov., D. javanensis sp. nov., and D. sumatrensis sp. nov. from Indonesia; D. wangi sp. nov. from East Malaysia (Sabah). Doryscus chujoi Takizawa, 1978 and D. varians (Gressitt & Kimoto, 1963) are removed from synonymy with D. testaceus Jacoby, 1887. Doryscus nigricollis Jiang, 1992 and D. marginicollis Jiang, 1992 are regarded as junior synonyms of D. varians (Gressitt & Kimoto, 1963). A lectotype is designated for Doryscus testaceus Jacoby, 1887.
A new silvanid genus Dentirotacorimus gen. nov. is described based on specimens collected from Ulu Gombak (Malay Peninsula), Malaysia. Two new species, D. reticulatus sp. nov. and D. zigzag sp. nov., are described herein. A key to species of this genus and a table listing states of diagnostic characters of Corimus-like genera, including this new genus, are provided.
The net-winged beetle genus Alyculus Kasantsev is reported from Peninsular Malaysia for the first time and a new species, A. malaypeninsularis sp. nov., is described and illustrated. An expanded identification key to Alyculus males is provided and the biology and distribution of the species are discussed.
The synchronous firefly genus Pteroptyx Olivier is reassessed from morphological, molecular, and habitat perspectives in Malaysia, and includes some reliably associated females described from morphological features and internal female reproductive anatomy. Phylogenetic analyses using combined morphological and molecular data (where available) for 158 taxa supported all the major features of the existing taxonomic categories within the Indopacific Luciolinae. They revealed a distinct Pteroptyx clade as a morphologically variable genus with Poluninius selangoriensis Ballantyne being newly synonymised with Luciola testacea Motschulsky, the type species, which is redescribed from the type series. Pteroptyx gelasina Ballantyne was shown to be distinct and three of the four morphological subdivisions within Pteroptyx malaccae (Gorham) considered useful. A new species Pt. balingiana Jusoh sp. nov. is described from Sarawak. A second specimen of Pt. gombakia Ballantyne is described and figured. Some females were reliably associated with identified males by molecular data, but investigation of their morphology showed consistent features that were for the most part not useful for species delineation, which still relies on association with the males and colour patterns. All females investigated had bursa plates.Habitat details for most Pteroptyx revealed an association with a riparian environment likely to support mangroves but not necessarily an obligatory association with mangroves or any particular species. Pteroptyx galbina Jusoh was found up to 30 km from the sea, and Pt. bearni Olivier displays in a variety of flowering plants alongside rivers, including mangroves.Keys to species and diagnoses of all species with coloured plates are given.
A worldwide review of the genus Callilanguria is presented. Three new species are described: C. weiweii Huang Yang, sp. nov. from Sabah, Malaysia; C. helleri Huang Yang, sp. nov. from Panay, the Philippines; and C. nigripes Huang Yang, sp. nov. from Samar Island, the Philippines. Callilanguria scrupulosa Heller, 1918 is transferred to the genus Doubledaya. The other species are C. eximia Fowler, 1885; C. gorhami Villiers, 1945; C. asymmetrica Heller, 1900; C. ruficeps Achard, 1923; C. milloti Villiers, 1945; C. stenosoma (Harold, 1879); C. flaviventris Fowler, 1886; C. wallacii Crotch, 1876; and C. luzonica Crotch, 1876. In total, twelve species are recognized in the genus Callilanguria, and a key to the described species of the genus is provided.
A new species of firefly (Coleoptera: Lampyridae) Pygoluciola dunguna Nada sp. nov. is described from males and reliably associated females, collected mainly from lowland dipterocarp forests of Peninsular Malaysia at elevations below 300 m a.s.l. This species is found to be terrestrial, flying at low heights between 10-15 m above ground in between the herbaceous plants. Males are intermediate between the two types of Pygoluciola Wittmer known until now. A list of species, and a key to males and known females of Pygoluciola is given. The female bursa has paired elongate hook like bursa plates.
In the present study we describe a new species of the genus Omophorus, O. (Sinomophorus) wallacei sp. nov., from Malaysian Borneo. Photographs of habitus and genitalia, a distribution map, and a key for the identification of Omophorus species are included. The definition of the subgenus Sinomophorus is revised and several new diagnostic characters are added. This is the first documented occurrence of Omophorus in Borneo, highlighting its potential diversity in the Oriental region and providing additional evidence of Ethiopian-Oriental disjunction within this genus.
This paper reports a new weevil genus Devernodes gen. n. established for five new species from Southeast Asia: D. alkippe sp. n. (China: Mt. Emei), D. asteria sp. n. (Vietnam: Tam Dao), D. chthonia sp. n. (Vietnam: Tam Dao; the type species), D. drimo sp. n. (Malaysia: Pasoh Forest Reserve) and D. methone sp. n. (Malaysia: Tanah Rata). All Devernodes are wingless and inhabit the forest leaf litter. Adult Devernodes share a combination of two head characters unique among weevils in Asia: antenna with apparently unsegmented club and 6-segmented antennal funicle, as well as strong constriction separating the eye-bearing rostrum from the head capsule. To test monophyly and investigate phylogenetic relationships of Devernodes, Maximum Likelihood phylogenetic analysis was undertaken using parts of mitochondrial (COI) and nuclear ribosomal (28S) genes, as well as the nuclear internal transcribed spacer (ITS2) from 14 Devernodes and 55 outgroup Curculionidae specimens. Results strongly corroborated monophyly of Devernodes and did not suggest its realistic sister-group. The new genus is assigned to the molytine tribe Lymantini (not represented in the DNA analysis) based on two potential synapomorphies: head markedly constricted behind eyes and presence of undivided female hemisternites IX (= "merged coxite and stylus"). Thus interpreted, Devernodes is the twelfth nominal genus of Lymantini and the first record of the tribe outside of the Americas. All original data (localities, DNA sequences, specimen images) are available online in public datasets dx.doi.org/10.5883/DS-DEVERNO1 and dx.doi.org/10.5883/DS-DEVERNO2.
Platiana Schimmel, 1993 is a moderately large genus in Dimini endemic to South East Asia. Hitherto only four species of this genus were known from Peninsular Malaysia. Here, we describe and figure Platiana cechovskyi sp. nov. from the Cameron Highlands. This species is characteristic by its relatively small body, dark coloration, and strongly serrate antennae which are rarely encountered in Dimini. An identification key to the Platiana species from Peninsular Malaysia as well as a distribution map for all known species are provided.
The pselaphine genus Sathytes Westwood is represented in Borneo by a single species. In this paper we describe four new species: S. borneoensis sp. nov., S. liuyei sp. nov., S. larifuga sp. nov., and S. shihongliangi sp. nov. All new species were collected by light traps in Sabah, and show unique combinations of male sexual characters that lead to ready species discriminations. A key to Sathytes species from Borneo is appended.
Twenty new species of Agrilus jewel beetles from the Oriental region are described and illustrated: Agrilus cicadelloides sp. nov. (Malaysia: Sarawak); A. draco sp. nov. (Malaysia: Sabah); A. hergovitsi sp. nov. (Malaysia: Johor); A. hik sp. nov.(Cambodia); A. ika sp. nov. (Solomon Islands); A. jankae sp. nov. (Indonesia: Sumatra, Singapore); A. jum sp. nov. (Laos); A. kon sp. nov. (Cambodia); A. mimicus sp. nov. (Laos); A. qom sp. nov. (Laos); A. titi sp. nov. (Malaysia: Perak); A. uxo sp. nov. (Vietnam); A. wos sp. nov. (Laos); A. xen sp. nov. (Laos); A. xia sp. nov. (Laos); A. xis sp. nov. (Laos); A. yoa sp. nov. (Laos, Vietnam); A. yuk sp. nov. (Laos); A. zao sp. nov. (Indonesia: Java and Sumba Islands); A. zim sp. nov. (Malaysia: Pahang).
Four new species of Grouvellinus Champion, 1923 with very long median pronotal carina are described from China: G. hongkongensis sp. nov. (Guangdong, Guangxi, Hong Kong), G. longiusculus sp. nov. (Jiangxi), G. mediocarinatus sp. nov. (Fujian, Guangdong), and G. robustus sp. nov. (Anhui). These species are obviously closely related to G. bishopi Jäch, 1984 described from Malaysia (Selangor). The latter is here recorded for the first time from Kedah and Perak (Malaysia). Habitus photographs, detailed line drawings of the male genitalia, as well as a key to the males of the species of Grouvellinus with very long median pronotal carina are provided.
The 'Cephennomicrus group' of Cephenniini includes nine genera distributed mostly in SE Asia (but with some species known also from other continents, except for the Americas). Four new Oriental species are described: Cephennococcus penangensis sp. n. (W Malaysia), Cephennomicrus matangensis sp. n. (E Malaysia), Cephennomicrus selangorensis sp. n. (E Malaysia), and Trurlia nova sp. n. (E Malaysia). Cephennomicrus selangorensis is unique in having the pronotum in both sexes with broad lateral impressions demarcated from the disc by a sharp ridge, a modification slightly similar to that of only one (Australian) congener.
The genus Carlschoenherria Bezděk, 2016 (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini) in the Philippines is reviewed. Three new species: Carlschoenherria adoradae Calcetas, new species, C. hadsallae Calcetas, new species, and C. gapudi Calcetas, new species are described, figured and compared with their close relatives. The following new combinations are proposed: Carlschoenherria palawana (Moser, 1915) new combination, C. philippinica (Brenske, 1894) new combination, C. argus (Burmeister, 1855) new combination, C. hastata (Arrow, 1938) new combination and C. vervex (Sharp, 1876) new combination. A new C. adoradae species-subgroup is proposed under the broader C. sulcipennis species-group. A key of the Carlschoenherria of the Philippines Islands, Malay Peninsula, Sumatra and Java is provided.
The genus Theopea Baly, 1864 is redefined. Seventeen species from Sundaland and the Philippines are recognized and classified into four species groups (including seven new species): T. impressa (Fabricius, 1801), T. longicollis (Jacoby, 1896), T. louwerensi Jolivet, 1951, T. lunduensis Mohamedsaid, 1998, T. chungi sp. nov. in the T. impressa group; T. flavipalpis Laboissère, 1940, T. guoi sp. nov., T. lui sp. nov., T. sabahensis sp. nov. in the T. flavipalpis group; T. elegantula Baly, 1864, T. pulchella Baly, 1864, T. fairmairei Duvivier, 1885, T. kedenburgi Weise, 1922, T. houjayi sp. nov., T. tsoui sp. nov., T. yuae sp. nov. in the T. pulchella group; and T. costata (Allard, 1889) in the T. costata group. The following new synonyms are established: Theopea pulchella Baly, 1864 = T. nigricollis Jacoby, 1892 syn. nov.; Theopea impressa (Fabricius, 1801) = T. impressa flavicornis Laboissère, 1940 syn. nov. and T. impressa malaccana Laboissère, 1940 syn. nov.; T. lunduensis Mohamedsaid, 1998 = T. sepilokensis Mohamedsaid, 2000 syn. nov. Lectotypes are designated for Crioceris impressa Fabricius, 1801, Ozomena longicollis Jacoby, 1896, Theopea elegantula Baly, 1864, T. fairmairei Duvivier, 1885, T. nigricollis Jacoby, 1892, and T. pulchella Baly, 1864. Theopea obliterata Jacoby, 1884, T. variabilis (Jacoby, 1887), T. incostata (Allard, 1889), T. clypeata Jacoby, 1896, T. modiglianii Jacoby, 1896, T. dohrni (Jacoby, 1899), T. viridipennis (Jacoby, 1899), and T. weberi (Weise, 1913) are removed from Theopea and regarded as species incertae sedis.
Megasybacodes brevitarsis Kakizoe, Maruyama Masumoto, a new genus and new species of the tribe Rhyparini, is described based on a single female from Borneo. It is allied to the genus Sybacodes Fairmaire, 1896, but easily distinguished by its short protarsi, broad flattened basal tarsomeres of the meso- and metatarsi, broad flattened tibiae, and wrinkled pronotal and elytral surfaces. Checklist of rhyparine species from Borneo is also provided.
The Agrilus purpurifrons species-group comprising twelve species from the Oriental region is defined and revised. A key to species is provided and complemented with illustrations of habitus and genitalia. Three new species are described: Agrilus cameronius sp. nov. (Malaysia); A. puncak sp. nov. (Indonesia); and A. vendibilis sp. nov. (Indonesia). The following taxonomic changes are proposed: the specific names lacroixi Obenberger, 1936 syn. nov. and chapaensis Descarpentries Villiers, 1967 syn. nov. are junior synonyms in the synonymy of A. morio Kerremans, 1895; the name rousselatae Baudon, 1968 stat. rev. is removed from the synonymy of A. lacroixi Obenberger, 1936 and revalidated as the specific name of A. rousselatae Baudon, 1968.