The Miltochrista obliquilinea (Swinhoe, 1901) species-group is revised. Four new species are described: M. konta Volynkin, Černý N. Singh, sp. n. (Thailand, Laos and Vietnam), M. adelfika Volynkin, N. Singh, Černý, Kirti Datta, sp. n. (India, Myanmar, China, Thailand, Laos and Vietnam), M. stenovalva Volynkin, N. Singh, Černý, Kirti Datta, sp. n. (India and Thailand) and M. lavides Volynkin, Černý N. Singh, sp. n. (Malaysia, Thailand, Myanmar, Laos and Vietnam). The lectotype for Lyclene obliquilinea Swinhoe, 1901 is designated. Adults, male and female genitalia are illustrated.
The genus Savarna Huber, 2005 comprises only five species, from southern Thailand, Peninsular Malaysia and Sumatra. In this study, five new species are described from Thailand: Savarna bannang sp. nov. (Yala), S. chiangmai sp. nov. (Chiangmai), S. huahin sp. nov. (Prachuap Kiri Khan), S. satun sp. nov. (Satun), S. thungsong sp. nov. (Nakhon Srithammarat). All new species are described from males and females. The distribution of S. chiangmai sp. nov. represent the northernmost record of the genus.
Betta nuluhon, new species, is described from a hill stream habitat in western Sabah. This species is allied to both B. chini and B. balunga, and differs from rest of its congeners in the B. akarensis group in having the following combination of characters: yellow iris when live; mature males with greenish-blue iridescence on opercle when live; mature fish with distinct transverse bars on caudal fin; slender body (body depth 22.1-25.2 % SL); belly area with faint reticulate pattern (scales posteriorly rimmed with black); absence of tiny black spots on anal fin; lateral scales 29-31 (mode 30); predorsal scales 20-21 (mode 20). Notes on a fresh series of B. chini are also provided.
Three new Indo-West Pacific species of pinnotherid crabs are described, one each of Arcotheres, Buergeres and Nepinnotheres. Arcotheres pollus, described from Paway Island, Mergui Archipelago, is most similar to A. boninensis (Stimpson, 1858), A. pernicola (Bürger, 1895) and A. purpureus (Alcock, 1900), sharing a transversely ovate carapace and long, slender, almost styliform dactyli of P4 and 5 that are about twice the length of those of P2 and 3. Buergeres choprai, described from Papua New Guinea, is most similar to B. deccanesis (Chopra, 1931) from eastern India but differentiated by segment proportions and setation of the walking legs. Buergeres tenuipes (Bürger, 1895) is synonymised with B. ortmanni (Bürger, 1895), which is also reported for the first time from Indonesia. A male of an undetermined species of Buergeres from the Philippines, possibly B. ortmanni, is figured and described, documenting the gonopod morphology in Buergeres for the first time. A key to the species of Buergeres based on females is provided. Nepinnotheres fulvia sp. nov. is also described from Papua New Guinea, and resembles N. cardii (Bürger, 1895) from the Philippines and Malaysia but can be distinguished by features of the chelipeds and maxilliped 3.
The first representative of the genus Cerapus in Malaysian waters, Cerapus bumbumiensis sp. nov. is described from specimens sampled from Pulau Bum Bum, Sabah (east Malaysia). The main identifying characteristics of this new amphipod species are: pereonites 1, 2 with constriction; male gnathopod 2 carpochelate, carpus large with long defining posterior tooth and well-developed anterodistal tooth; pereopod 6 coxa with fine hair/fringe setae ventrally; and telson with deep cleft. An updated identification key for the 23 known species in the genus is also presented.
A new species of Clidicini ant-like stone beetles, Clidicus mawarensis sp. n., is described and illustrated. The holotype male was collected in East Malaysia (Borneo: Sabah); the new species belongs to a group of large-bodied Clidicus, and shows similarities to C. ganglbaueri Reitter; the male has unusually complex structures of the aedeagal apical region.
Pygoluciola dunguna Nada, 2018 was described from Peninsular Malaysia, using males and reliably associated females. This paper details description of the larva which has been conclusively identified as Pygoluciola dunguna based on DNA barcoding technique and uses morphology, brief habitat and behavioural data. A total of 70 larval specimens were measured and their main features described. The larvae exhibit a riparian or semi-aquatic behaviour, observed crawling on the sandy edge of shallow streams. The stake-like projections along the length of the body suggest a form of defensive mechanism from falling prey to aquatic predators.
Cyana Walker, 1854 is one of the most species-rich Erebiidae genera within the tribe Lithosiini Billberg of the subfamily Arctiinae Leach. The genus is widespread from Sub-Saharan Africa and Madagascar through southern and eastern Asia to New Guinea and Australia with a diversity hot spot in South East Asia. A striking species with contrasting red and orange wing pattern, C. bellissima (Moore, 1878) was described from northern India and recorded from the Himalayas, China and Indochina (Fang 2000; Černý Pinratana 2009; Singh et al. 2020). Another closely related species, C. stresemanni (Rothschild, 1936) (= bellissima inouei Kishida, 1993) is distributed in the Peninsular Malaysia (Rothschild 1936; Kishida 1993; Bucsek 2012). During examination of extensive unsorted Lithosiini materials housed in the MWM/ZSM and the private collection of the senior author, a series of peculiar specimens from southern Vietnam provisionally identified as 'C. bellissima' was found. These specimens, however, display certain external differences from other populations of C. bellissima and C. stresemanni, suggesting the presence of a further taxon related to C. bellissima. The examination of the male and the female genitalia of the southern Vietnamese specimens has confirmed their specific distinctness and they are described in this paper as a new species.
We found a uniquely colored dicroglossid frog of the genus Occidozyga from western Sarawak, East Malaysia. It is divergent from other congeners in morphology and mtDNA sequences. In a molecular phylogeny, this species is the sister lineage to the continental species O. lima and O. martensii with weak support. The species is small with SVL 16-18 mm in males and 18-19 mm in females, without dorsolateral fold but with transverse wrinkles on dorsum, tips of fingers lacking disks but of toes with disks, only first and second toes webbed to disks, and orange-brown dorsum with dark brown band. We thus describe it as a new species.
Tanyxiphium longissimum Huber, syn. n. (Hymenoptera: Mymaridae) is synonymized under T. harriet (Zeya) based on examination of specimens from Hainan Island and Yunnan Province in the Oriental part of China, Sulawesi Island in Indonesia, Peninsular Malaysia, and Thailand. These are new country distribution records except for Thailand. The previously unknown male of T. harriet is described from Sulawesi Island, and the female is redescribed and illustrated based on non-type material examined.
A new, diminutive species of Rock Gecko Cnemaspis tubaensis sp. nov. of the C. kumpoli group, is described from Tuba Island, Langkawi Archipelago, Kedah, Peninsular Malaysia and is differentiated from all other species in the kumpoli group by having a unique combination of morphological and color pattern characteristics, including a maximum SVL of 37.0 mm; 10 or 11 supralabials; eight or nine infralabials; 15-18 semi-linearly arranged paravertebral tubercles; lateral caudal furrow present; lateral caudal tubercles on the anterior portion of the tail; caudal tubercles not encircling tail; five or six precloacal pores; 28 or 29 subdigital lamellae on the fourth toe; smooth ventrals; smooth subcaudals with an enlarged median row of scales; subcaudal region light-grey and speckled with yellow; absence of light-colored ocelli on the shoulder; no yellow postscapular band; dorsum light-brown with sage-green blotches and black spots; flanks with scattered yellow spots; absence of black gular markings in both sexes; and 13.0-22.0% uncorrected pairwise sequence divergences in the NADH dehydrogenase subunit 2 gene (ND2). Cnemaspis tubaensis sp. nov. is the fourth species of Cnemaspis to be described from the Langkawi Archipelago and underscores the underestimated biodiversity of the islands which is in need of more thorough herpetological inventories.
Megalogomphus sumatranus (Krüger, 1899) and its allies in Sundaland are reviewed. The accessory genitalia of the males of this genus, hardly considered previously, are found to be taxonomically informative. The taxon from Borneo previously treated as M. sumatranus is described from both sexes as M. buddi sp. nov. (holotype ♂ Sungai Datai, Nanga Bloh, Lanjak Entimau Wildlife Sanctuary, Kapit Division, Sarawak, Malaysia, 22 viii 2013, leg. J. anak Awan M. anak Adau; deposited at the Naturalis Biodiversity Center, Leiden, the Netherlands). Megalogomphus borneensis (Laidlaw, 1914), described as a subspecies of M. icterops (Martin, 1903) and subsequently relegated to the synonymy of that species, is considered to be a distinct species. Megalogomphus icterops is however considered to be a junior synonym of M. sumatranus. A re-description of the holotype of Megalogomphus borneensis is provided as is the first description of the female. Descriptive notes with illustrations of Megalogomphus sumatranus are given.
An overview of the Oriental species of the nominate subgenus of Camptopteroides Viggiani (Hymenoptera: Mymaridae) is given. Two new species, C. (Camptopteroides) formosa Manickavasagam Sankararaman sp. n. from India and C. (Camptopteroides) reducta Triapitsyn sp. n. from Thailand and Malaysia, are described, and C. formosa additionally compared to two unnamed species. The holotype male of the type species of this genus, C. armata Viggiani from Sri Lanka, is diagnosed and illustrated. A key to Old World species of Camptopteroides is provided.
The genus Olios Walckenaer, 1837 is revised, a generic diagnosis is given and an identification key to eight species groups is provided. Olios in its revised sense includes 87 species and is distributed in Africa, southern Europe and Asia. Three species groups are revised in this first part, an identification key to species for each group is provided, five new species are described and all included species are illustrated. The Olios argelasius-group includes O. argelasius Walckenaer, 1806, O. canariensis (Lucas, 1838), O. pictus (Simon, 1885), O. fasciculatus Simon, 1880 and O. kunzi spec. nov. (male, female; Namibia, Zambia, South Africa); it is distributed in the Mediterranean region, northern Africa including Canary Islands, in the Middle East, South Sudan, East Africa, and southern Africa. The Olios coenobitus-group includes O. angolensis spec. nov. (male; Angola), O. coenobitus Fage, 1926, O. denticulus spec. nov. (male; Java), O. erraticus Fage, 1926, O. gambiensis spec. nov. (male, female; Gambia), O. milleti (Pocock, 1901b), O. mordax (O. Pickard-Cambridge, 1899) and O. pusillus Simon, 1880; it is distributed in Africa (Gambia, Angola, Tanzania, Madagascar) and Asia (India, Sri Lanka, Indonesia: Java). The Olios auricomis-group includes only O. auricomis (Simon, 1880), distributed in Africa south of 10°N. Other species groups are introduced briefly and will be revised in forthcoming revisions. The Olios correvoni-group includes currently O. claviger (Pocock, 1901a), O. correvoni Lessert, 1921, O. correvoni choupangensis Lessert, 1936, O. darlingi (Pocock, 1901a), O. faesi Lessert, 1933, O. freyi Lessert, 1929, O. kassenjicola Strand, 1916b, O. kruegeri (Simon, 1897a), O. quadrispilotus (Simon, 1880) comb. nov., O. lucieni comb. nov. nom. nov., O. sjostedti Lessert, 1921 and O. triarmatus Lessert, 1936; it is distributed in Africa (Zimbabwe, Tanzania incl. Zanzibar, Angola, Congo, Central Africa, South Africa, Botswana; O. darlingi was recorded from Zimbabwe and Botswana and not from South Africa). The Olios rossettii-group includes: O. baulnyi (Simon, 1874), O. bhattacharjeei (Saha Raychaudhuri, 2007), O. brachycephalus Lawrence, 1938, O. floweri Lessert, 1921, O. jaldaparaensis Saha Raychaudhuri, 2007, O. japonicus Jäger Ono, 2000, O. kolosvaryi (Caporiacco, 1947b) comb. nov., O. longipes (Simon, 1884b), O. lutescens (Thorell, 1894), O. mahabangkawitus Barrion Litsinger, 1995, O. obesulus (Pocock, 1901b), O. rossettii (Leardi, 1901), O. rotundiceps (Pocock, 1901b), O. sericeus (Kroneberg, 1875), O. sherwoodi Lessert, 1929, O. suavis (O. Pickard-Cambridge, 1876), O. tarandus (Simon, 1897d), O. tener (Thorell, 1891) and O. tiantongensis (Zhang Kim, 1996); it is distributed in the Mediterranean region, in Africa (especially eastern half) and Asia (Middle East and Central Asia to Japan, Philippines and Java). The Olios nentwigi-group includes O. diao Jäger, 2012, O. digitatus Sun, Li Zhang, 2011, O. jaenicke Jäger, 2012, O. muang Jäger, 2012, O. nanningensis (Hu Ru, 1988), O. nentwigi spec. nov. (male, female; Indonesia: Krakatau), O. perezi Barrion Litsinger, 1995, O. scalptor Jäger Ono, 2001 and O. suung Jäger, 2012; it is distributed in Asia (Thailand, Laos, Vietnam, Cambodia, China, Taiwan, Indonesia, Philippines), Papua New Guinea and Mariana Islands. Olios diao is newly recorded from Cambodia and Champasak Province in Laos. The Olios stimulator-group includes O. admiratus (Pocock, 1901b), O. hampsoni (Pocock, 1901b), O. lamarcki (Latreille, 1806) and O. stimulator Simon, 1897c; it is distributed in Africa (Madagascar, Seychelles), Middle East and South Asia (United Arab Emirates, Iraq, Afghanistan, Pakistan, India, Maldives, Sri Lanka). The Olios hirtus-group includes O. bungarensis Strand, 1913b, O. debalae (Biswas Roy, 2005), O. ferox (Thorell, 1892), O. hirtus (Karsch, 1879a), O. igraya (Barrion Litsinger, 1995) comb. nov., O. menghaiensis (Wang Zhang, 1990), O. nigrifrons (Simon, 1897b), O. punctipes Simon, 1884a, O. punctipes sordidatus (Thorell, 1895), O. pyrozonis (Pocock, 1901b), O. sungaya (Barrion Litsinger, 1995) comb. nov., O. taprobanicus Strand, 1913b and O. tikaderi Kundu et al., 1999; it is distributed in South, East and Southeast Asia (Sri Lanka, India, Nepal, Bangladesh, Myanmar, China, Laos, Thailand, Cambodia, Vietnam, Malaysia, Indonesia, Philippines). Nineteen synonyms are recognised: Nisueta Simon, 1880, Nonianus Simon, 1885, both = Olios syn. nov.; O. spenceri Pocock, 1896, O. werneri (Simon, 1906a), O. albertius Strand, 1913a, O. banananus Strand, 1916a, O. aristophanei Lessert, 1936, all = O. fasciculatus; O. subpusillus Strand, 1907c = O. pusillus; O. schonlandi (Pocock, 1900b), O. rufilatus Pocock, 1900c, O. chiracanthiformis Strand, 1906, O. ituricus Strand, 1913a, O. isongonis Strand, 1915, O. flavescens Caporiacco, 1941 comb. nov., O. pacifer Lessert, 1921, all = O. auricomis; Olios sanguinifrons (Simon, 1906b) = O. rossettii Leardi, 1901; O. phipsoni (Pocock, 1899), Sparassus iranii (Pocock, 1901b), both = O. stimulator; O. fuligineus (Pocock, 1901b) = O. hampsoni. Nine species are transferred to Olios: O. gaujoni (Simon, 1897b) comb. nov., O. pictus comb. nov., O. unilateralis (Strand, 1908b) comb. nov. (all three from Nonianus), O. affinis (Strand, 1906) comb. nov., O. flavescens Caporiacco, 1941 comb. nov., O. quadrispilotus comb. nov., O. similis (Berland, 1922) comb. nov. (all four from Nisueta), O. sungaya (Barrion Litsinger, 1995) comb. nov., O. igraya (Barrion Litsinger, 1995) comb. nov. (both from Isopeda L. Koch 1875). Olios lucieni nom. nov. comb. nov. is proposed for Nisueta similis Berland, 1922, which becomes a secondary homonym. The male of O. quadrispilotus comb. nov. is described for the first time. Sixteen species are currently without affiliation to one of the eight species groups: O. acolastus (Thorell, 1890), O. alluaudi Simon, 1887a, O. batesi (Pocock, 1900c), O. bhavnagarensis Sethi Tikader, 1988, O. croseiceps (Pocock, 1898b), O. durlaviae Biswas Raychaudhuri, 2005, O. gentilis (Karsch, 1879b), O. gravelyi Sethi Tikader, 1988, O. greeni (Pocock, 1901b), O. inaequipes (Simon 1890), O. punjabensis Dyal, 1935, O. ruwenzoricus Strand, 1913a, O. senilis Simon, 1880, O. somalicus Caporiacco, 1940, O. wroughtoni (Simon, 1897c) and O. zulu Simon, 1880. Five of these species are illustrated in order to allow identification of the opposite (male) sex and to settle their systematic placement. Thirty-seven species are considered nomina dubia, mostly because they were described from immatures, three of them are illustrated: O. abnormis (Blackwall, 1866), O. affinis (Strand, 1906) comb. nov., O. africanus (Karsch, 1878), O. amanensis Strand, 1907a, O. annandalei (Simon, 1901), O. bivittatus Roewer, 1951, O. ceylonicus (Leardi, 1902), O. conspersipes (Thorell, 1899), Palystes derasus (C.L. Koch, 1845) comb. nov., O. detritus (C.L. Koch, 1845), O. digitalis Eydoux Souleyet, 1842, O. exterritorialis Strand, 1907b, O. flavovittatus (Caporiacco, 1935), O. fugax (O. Pickard-Cambridge, 1885), O. guineibius Strand, 1911c, O. guttipes (Simon, 1897a), O. kiranae Sethi Tikader, 1988, O. longespinus Caporiacco, 1947b, O. maculinotatus Strand, 1909, O. morbillosus (MacLeay, 1827), O. occidentalis (Karsch, 1879b), O. ornatus (Thorell, 1877), O. pagurus Walckenaer, 1837, O. patagiatus (Simon, 1897b), O. praecinctus (L. Koch, 1865), O. provocator Walckenaer, 1837, O. quesitio Moradmand, 2013, O. quinquelineatus Taczanowski, 1872, O. sexpunctatus Caporiacco, 1947a, Heteropoda similaris (Rainbow, 1898) comb. rev., O. socotranus (Pocock, 1903), O. striatus (Blackwall, 1867), O. timidus (O. Pickard-Cambridge, 1885), Remmius variatus (Thorell, 1899) comb. nov., O. vittifemur Strand, 1916b, O. wolfi Strand, 1911a and O. zebra (Thorell, 1881). Eighty-nine species are misplaced in Olios but cannot be affiliated to any of the known genera. They belong to the subfamilies Deleninae Hogg, 1903, Sparassinae Bertkau, 1872 and Palystinae Simon, 1897a, nineteen of them are illustrated: O. acostae Schenkel, 1953, O. actaeon (Pocock, 1898c), O. artemis Hogg, 1915, O. atomarius Simon, 1880, O. attractus Petrunkevitch, 1911, O. auranticus Mello-Leitão, 1918, O. benitensis (Pocock, 1900c), O. berlandi Roewer, 1951, O. biarmatus Lessert, 1925, O. canalae Berland, 1924, O. caprinus Mello-Leitão, 1918, O. chelifer Lawrence, 1937, O. chubbi Lessert, 1923, O. clarus (Keyserling, 1880), O. coccineiventris (Simon, 1880), O. corallinus Schmidt, 1971, O. crassus Banks, 1909, O. debilipes Mello-Leitão, 1945, O. discolorichelis Caporiacco, 1947a, O. erroneus O. Pickard-Cambridge, 1890, O. extensus Berland, 1924, O. fasciiventris Simon, 1880 , O. feldmanni Strand, 1915, O. fimbriatus Chrysanthus, 1965, O. flavens Nicolet, 1849, O. fonticola (Pocock, 1902), O. formosus Banks, 1929, O. francoisi (Simon, 1898a), O. fulvithorax Berland, 1924, O. galapagoensis Banks, 1902, O. gaujoni (Simon, 1897b) comb. nov., O. giganteus Keyserling, 1884, O. hoplites Caporiacco, 1941, O. humboldtianus Berland, 1924, O. insignifer Chrysanthus, 1965, O. insulanus (Thorell, 1881), O. keyserlingi (Simon, 1880), O. lacticolor Lawrence, 1952, O. lepidus Vellard, 1924, O. longipedatus Roewer, 1951, O. machadoi Lawrence, 1952, O. macroepigynus Soares, 1944, O. maculatus Blackwall, 1862, O. marshalli (Pocock, 1898a), O. mathani (Simon, 1880), O. minensis Mello-Leitão, 1917, O. monticola Berland, 1924, O. mutabilis Mello-Leitão, 1917, O. mygalinus Doleschall, 1857, O. mygalinus cinctipes Merian, 1911, O. mygalinus nirgripalpis Merian, 1911, O. neocaledonicus Berland, 1924, O. nigristernis (Simon, 1880), O. nigriventris Taczanowski, 1872, O. oberzelleri Kritscher, 1966, O. obscurus (Keyserling, 1880), O. obtusus F.O. Pickard-Cambridge, 1900, O. orchiticus Mello-Leitão, 1930, O. oubatchensis Berland, 1924, O. paraensis (Keyserling, 1880), O. pellucidus (Keyserling, 1880), O. peruvianus Roewer, 1951, O. pictitarsis Simon, 1880, O. plumipes Mello-Leitão, 1937, O. princeps Hogg, 1914, O. pulchripes (Thorell, 1899), O. puniceus (Simon, 1880), O. roeweri Caporiacco, 1955a, O. rubripes Taczanowski, 1872, O. rubriventris (Thorell, 1881), O. rufus Keyserling, 1880, O. sanctivincenti (Simon, 1898b), O. similis (O. Pickard-Cambridge, 1890), O. simoni (O. Pickard-Cambridge, 1890), O. skwarrae Roewer, 1933, O. spinipalpis (Pocock, 1901a), O. stictopus (Pocock, 1898a), O. strandi Kolosváry, 1934, O. subadultus Mello-Leitão, 1930, O. sulphuratus (Thorell, 1899), O. sylvaticus (Blackwall, 1862), O. tamerlani Roewer, 1951, O. tigrinus (Keyserling, 1880), O. trifurcatus (Pocock, 1900c), O. trinitatis Strand, 1916a, O. velox (Simon, 1880), O. ventrosus Nicolet, 1849, O. vitiosus Vellard, 1924 and O. yucatanus Chamberlin, 1925. Seventeen taxa are transferred from Olios to other genera within Sparassidae, eight of them are illustrated: Adcatomus luteus (Keyserling, 1880) comb. nov., Eusparassus flavidus (O. Pickard-Cambridge, 1885) comb. nov., Palystes derasus (C.L. Koch, 1845) comb. nov., Heteropoda similaris (Rainbow, 1898) comb. rev., Remmius variatus (Thorell, 1899) comb. nov., Nolavia audax (Banks, 1909) comb. nov., Nolavia antiguensis (Keyserling, 1880) comb. nov., Nolavia antiguensis columbiensis (Schmidt, 1971) comb. nov., Nolavia fuhrmanni (Strand, 1914) comb. nov., Nolavia helva (Keyserling, 1880) comb. nov., Nolavia stylifer (F.O. Pickard-Cambridge, 1900) comb. nov., Nolavia valenciae (Strand, 1916a) comb. nov., Nungara cayana (Taczanowski, 1872) comb. nov., Polybetes bombilius (F.O. Pickard-Cambridge, 1899) comb. nov., Polybetes fasciatus (Keyserling, 1880) comb. nov., Polybetes hyeroglyphicus (Mello-Leitão, 1918) comb. nov. and Prychia paalonga (Barrion Litsinger, 1995) comb. nov. One species is transferred from Olios to the family Clubionidae Wagner, 1887: Clubiona paenuliformis (Strand, 1916a) comb. nov.
Coeliccia junis sp. nov. (holotype male from Borneo, Sarawak, Bintulu Division, Planted Forest Project, Bukit Mina Wildlife Corridor, "Day 4" stream near Bukit Nyegoh and Bukit Jugam, near small brown water pool, 10 viii 2018, deposited in the Naturalis Biodiversity Center, Leiden, the Netherlands) and Coeliccia roberti sp. nov. (holotype male from Borneo, Sarawak, Miri Division, Lambir Hills National Park, small stream on Oil Well Trail, 22 iv 2011, deposited in the Naturalis Biodiversity Center, Leiden, the Netherlands) are described from Borneo. Both new species belong to the Coeliccia membranipes-group and bring the number of named species known from the group from Borneo to nine and the total number of named species from Borneo currently placed in Coeliccia to 15. Coeliccia junis is only known from a small area in Sarawak, C. roberti is also known from Brunei. Both species are considered likely to be closely allied with C. macrostigma Laidlaw.
Six new species belonging to Belisana Thorell, 1898 are described from Southeast Asia: Belisana bachma sp. nov. (Vietnam; male, female), B. cucphuong sp. nov. (Vietnam; male, female), B. jaegeri sp. nov. (Malaysia; male, female), B. kachin sp. nov. (Myanmar; male, female), B. putao sp. nov. (Myanmar; male) and B. tarang sp. nov. (Indonesia; male, female). These new species bring the total number of Belisana to 143 species worldwide.
The genus Surenus Distant, 1901 (Hemiptera: Heteroptera: Pentatomoidea: Pentatomidae: Pentatominae: Halyini) is revisited and found to be a junior subjective synonym of the genus Agathocles Stål, 1876 (currently Pentatominae: Rolstoniellini). The genus Agathocles and its type species, Agathocles limbatus Stål, 1876, are redescribed. Lectotype of Surenus normalis Distant, 1901 (= Agathocles normalis (Distant, 1901) comb. nov.) is designated and the unknown male of the species is described. Agathocles yunnanensis Zhang Lin, 1984, syn. nov., is considered junior subjective synonym of A. limbatus. Two new species are described: Agathocles flavipes sp. nov. from India (Andhra Pradesh, Karnataka, Maharashtra, and Tamil Nadu) and A. joceliae sp. nov. from Malaysia (Kelantan, Perak). The new species differ from their congeners mainly by the morphology of mandibular plates, length of antennomeres I, IIa and IIb, body length, and structure of male genitalia. Agathocles dubius Distant, 1921 is transferred to the genus Caystrus Stål, 1861 (Pentatominae: Caystrini) based on examination of its holotype with the resulting new combination: Caystrus dubius (Distant, 1921), comb. nov. One new combination is proposed, Paramecocoris ruficornis (Fieber, 1851), comb. nov. (from preoccupied Paramecus Fieber, 1851), and its type locality is clarified as Tenasserim (south Myanmar). Gender agreement and authorship of the name Riazocoris niger Ahmad Afzal, 1977 in Ahmad et al. (1977: 161) are corrected and status of its name bearing type is clarified as lectotype. The following new distribution records are given: A. limbatus from Cambodia, China (Guangxi, Tibet), Laos and Thailand, A. normalis, Caystrus obscurus (Distant, 1901a) and Critheus lineatifrons Stål, 1869 from Laos, Amasenus corticalis Stål, 1863 from Cambodia, Indonesia (E Kalimantan), Laos, Myanmar and Thailand, and Rolstoniellus boutanicus (Dallas, 1849) from Vietnam. Based on characters of external morphology and genitalia, the genus Agathocles is compared with representatives of the genera Halys Fabricius, 1803 (Halyini), Caystrus (Caystrini), Laprius Stål, 1861 (Myrocheini), and Exithemus Distant, 1902 (currently in Rolstoniellini). As a result, the genus Agathocles is here transferred to the tribe Caystrini. The genus Kyrtalus Van Duzee, 1929 is tentatively placed in Myrocheini based on the presence of sulcate mesosternum and femora provided with teeth.
Six new species of the cephenniine genus Hlavaciellus Jałoszyński are described: H. cincinnalis sp. n. (Malaysia: Sabah); H. carinatus sp. n. (Malaysia: Pahang); H. microtuberculatus sp. n. (Malaysia: Pahang); H. diversipilosus sp. n. (Malaysia: Pahang); H. concavus sp. n. (Indonesia: Sumatra); and H. sumatranus sp. n. (Indonesia: Sumatra). An updated key to identification of males of all nominal species of Hlavaciellus is given.
Although the first issue of Zootaxa appeared in 2001 it was not until two years later, in August 2003, that this aspiring and inspiring new journal issued the first paper on the insect Order Thysanoptera, in Volume 268. Moreover, it was not until February 2005 that the second paper concerning this group appeared in Zootaxa. The subsequent expansion is summarized most succinctly by the number of Thysanoptera papers that appeared in Zootaxa in each of the four five-year periods of these two decades: 5; 40; 92; 134 (see Table 1). The 270 papers concerning this group of insects that appeared in Zootaxa over the 20-year period involved more than 120 authors. These papers were submitted by workers from about 30 different countries, but most of them were from areas of high but previously unexplored species diversity, particularly Australia, Brazil, China, India, Iran, Japan and Malaysia. However, significant contributions were submitted from the far north, including Poland and Russia, as well as the far south, such as Argentina and New Zealand. One reason for the popularity of Zootaxa amongst workers on thrips is presumably the knowledge that this section is edited by two active students of these insects. The editors are pleased to have rejected no more than five papers over these two decades, but they provide much help to authors in shaping manuscripts to ensure that the submitted information is both appropriate, scientifically correct, novel and clearly expressed. Moreover, the journal ensures that manuscripts are published very quickly, usually within four weeks of acceptance by the editors. For the readers a further advantage of Zootaxa is that just over 50% of the published articles on Thysanoptera are freely available on the web, as authors have arranged for Open Access. The thrips publications issued in Zootaxa have included descriptions of 563 new species and 41 new genera of Thysanoptera. These new species represent 9% of the 6300 valid extant species currently listed in this Order, and the new genera represent 5% of the 780 currently recognized genera (ThripsWiki 2021). Many of the publications are only of one or two pages and are issued as Correspondence. Each of these deals with a single new species, or a previously unknown male of a species, or some new and particularly unusual record for a country or host plant. At the opposite extreme are the Monographs that involve revisions of all of the species in a genus, such as the 60 species recognised in the South American genus Holopothrips, or the 30 species known in the worldwide genus Mycterothrips. Others provide illustrated keys to large numbers of genera, such as the 100 genera of Phlaeothripinae that have been recorded from South East Asia. Such extensive studies provide the factual resource on which many of the Articles published in Zootaxa are based. These Articles range from taxonomic revisions of small genera, or of the species found in particular geographic areas, through studies on character state variation and homologies, to historical accounts and catalogues. The very considerable increase in information in recent years about the taxonomic and biological diversity of this group of insects (Mound Hastenpflug-Vesmanis2021) owes much to the existence of the journal Zootaxa.
The final instar larva of Orthetrum borneense Kimmins, 1936, is described and figured for the first time based on exuviae from three male and six female larvae collected in Sarawak, Borneo (East Malaysia). It is compared with an early instar larva, which was matched to the adult O. borneense by DNA barcoding, and the known larvae of other species of this genus that occur in the region.