1 Ex-Situ Conservation Division, Department of Wildlife and National Parks (DWNP), Peninsular Malaysia, Ministry of Natural Resources and Environment Malaysia (NRE), KM 10 Jalan Cheras, 56100 Kuala Lumpur, Malaysia ; Department of Zoology, Faculty of Resource Science and Technology, Universiti Malaysia Sarawak (UNIMAS), 94300 Kota Samarahan, Sarawak, Malaysia